IRIG: PGC-1/Akt interaction in "Nature" and PPARg regulator in "Cell"

Liver produces glucose from amino acids through gluconeogenesis. If glucose is too much to be consumed by the peripheral tissues (muscle or fat), liver will covert the carbohydrate into fatty acids or triglycerides through hepatic de novo lipogenesis, resulting in an increase in plasma triglyceride and a reduction in plasma high-density lipoprotein. This is demonstrated in human in a new study in PNAS (See attachment 1). The hepatic gluconeogenesis and lipogenesis is regulated by insulin, which inhibits the two processes through the PI3K/Akt signaling pathway. In a “Nature” paper, a new event at downstream Akt is identified for the insulin-mediated inhibition. Akt2 was found to phosphorylate PGC-1 resulting in inhibition of PGC-1 function (See attachment 2). This event leads to shutdown of gene expression in the programs for gluconeogenesis and lipogenesis in the liver. PGC-1 is a coactivator of PPARg. In a “Cell” paper, a small molecule (Harmine) from a botanical product was found to increase insulin sensitivity by inducing PPARg gene expression. Harmine acts by inhibition of the Wnt signaling pathway (See attachment 3, 2MB in size).
 
The paper in attachment 1 is recommended by Dr. Zhenqi Liu (University of Virginia School of medicine), and paper in attachment 3 is recommended by Dr. Eugene Chen (University of Michigan Medical Center).
 
Have a nice weekend.
        
By Jianping at PBRC

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Jianping Ye, MD

Professor of Molecular Biology
Pennington Biomedical Research Center
Louisiana State University System
6400 Perkins Road
Baton Rouge, LA 70808
Phone: (225)763-3163
E-mail: yej@pbrc.edu
Webpage: http://labs.pbrc.edu/generegulation/index.htm
 
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RIG: Akt review in Cell

Akt (PKB) is a serine kinase in the signaling pathway of insulin receptor. In the metabolic branch of the signaling pathway, Akt mediates insulin signal for GLUT4 translocation, glycogen synthesis (Through GSK-3) and protein synthesis (through mTOR/S6K). However, this is only a part of biological activities of Akt in cells. A recent review in Cell gives a nice update on molecules at up- and down-stream of Akt. By the way, a figure in "Cell Signaling" website is helpful to get a full picture of Akt activities. See attached PDF files.
 
        In the current issue of "Molecular Cell", a new study made a progress in the signaling pathway of FFA. It is known that FFA may induce insulin resistance by targeting IRS-1 in the insulin receptor pathway. This activity is dependent on activation of PKC-JNK kinase axis by FFA. However, the molecules linking PKC to JNK are not completely identified in the FFA pathway. This study shows that a kinase by name of MLK (mixed lineage kinase) mediates PKC signal for JNK activation. Knockout of MLK may reduce Ser307 phosphorylation in IRS-1.
See attached PDF file.
 
by Jianping at PBRC

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Jianping Ye, MD

Professor of Molecular Biology
Pennington Biomedical Research Center
Louisiana State University System
6400 Perkins Road
Baton Rouge, LA 70808
Phone: (225)763-3163
E-mail: yej@pbrc.edu
Webpage: http://labs.pbrc.edu/generegulation/index.htm
     
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IRIG: Longevity in "Science" and Lipid-ER interaction in "Nature"

Insulin resistance in obesity increases risk for type 2 diabetes and cardiovascular diseases. These diseases usually accelerate aging and reduce lifespan. However, a study in the current issue of Science suggests that insulin resistance from IRS-2 knockout increased lifespan together with body weight. The abstract states that "In mice, less insulin receptor substrate–2 (Irs2) signaling throughout the body or just in the brain extended life span up to 18%." The study suggests that the role of insulin resistance in aging and longevity of obese subjects is not that simple. See attached paper that is recommended by Dr. Dixit at PBRC.
       
         ER stress in obesity has been used to explain insulin resistance. Oversupply of lipid may cause ER stress. However, it is not clear how lipid induces ER stress in obese condition. In the current issue of Nature, an article presents a hypothesis for the regulation of ER function by lipid.
See attached paper.
 
By Jianping at PBRC

------------------------------------------------------          


Jianping Ye, MD

Professor of Molecular Biology
Pennington Biomedical Research Center
Louisiana State University System
6400 Perkins Road
Baton Rouge, LA 70808
Phone: (225)763-3163
E-mail: yej@pbrc.edu
Webpage: http://labs.pbrc.edu/generegulation/index.htm
      
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